(Deciduous) Trees. Leaves spiral, imparipinnate, petioled. Leaflets subopposite or opposite, entire, mostly with internerval veins. Inflorescences paniculate, axillary and/or terminal. Flowers unisexual (plants dioecious). Calyx 4-lobed, greatly enlarged in fruit. Petals 4, imbricate, glabrous or sparsely hairy on the outer surface. Stamens 4, filaments long, often thin, glabrous; anthers usually ovoid, rarely oblong, dorsifixed or dorso-basifixed, abortive in ♀. Disk intrastaminal, hairy, round or slightly 4-angular, flat or discoid, 4-notched or-lobed; or pulvinate and 4-grooved. Ovary 1-celled, densely hairy; style 3-(rarely 4-)lobed; stigmas 3 (rarely 4). Sterile pistil in ♂ very small. Drupe 1-celled, densely brown hairy; subtended by the enlarged calyx, the 4 lobes wing-like; endocarp cartilaginous. Seed with testa adhering to the endocarp; embryo straight, cotyledons free, plano-convex.
Usually in dryland forest, also on inundated riverbanks and in freshwater swamps, in the lowland, rarely higher from 600-1450 m.
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