Trees, dioecious. Leaves paripinnate, alternate, estipulate, axis carinate abaxially, flat adaxially, triangular in transverse section; leaflets usually many jugate, entire or obtusely dentate. Thyrses axillary near apex, solitary or several fascicled; bracts and bracteoles small. Flowers unisexual, actinomorphic. Calyx shallowly cupular, deeply 5-lobed; sepals ovate or triangular, imbricate. Petals 5, ovate or subovate, with 1 large scale adaxially, scale thick, apex reflexed, abaxial surface and margin hirsute. Disk deeply cupular, slightly fleshy, margin thin, deeply wavy. Stamens (male flowers) 8, sometimes 7; filaments filiform, densely hairy; anthers broadly ovoid or ellipsoid-ovoid, connectives prominent and glandlike. Ovary (female flowers) 3-loculed; ovule 1 per locule, campylotropous; style terminal, longer than ovary; stigma inconspicuously 3-lobed. Capsules loculicidal into 3 schizocarps, 3-loculed or 1 or 2 small and sterile (without seed). Seeds 1-3; testa brown, shiny, leathery; hilum transversely elliptic, large, broad.