Herbs perennial, dioecious. Stems erect, simple, winged or sharply, longitudinally angled, glabrous. Leaves alternate; stipules narrowly triangular or narrowly lanceolate, 6-9 × 1.5-2 mm; petiole 3-10 mm; leaf blade herbaceous, obliquely obovate or elliptic, 8-18 × 4.4-8.3 cm, venation pinnate, major lateral veins ill-defined, broader outer half rounded or auriculate, margin undulate-crenate, apex acuminate or caudate-acuminate; cystoliths obscure, sparse; nanophylls 2-15 mm. Staminate inflorescences 2.5-3.5 cm in diam., peduncle 3.5-5.0 cm, pubescent; staminate flowers: tepals 5, elliptic; stamens 5; rudimentary pistil very small. Pistillate inflorescences ca. 1.1 cm in diam., peduncle 0.6-1.2 cm; pistillate flowers: tepals 5. Achenes ellipsoidal, tuberculate. Fl. Sep-Dec.