Herbs perennial, monoecious or dioecious. Stems erect or ascending, usually simple, densely strigose. Leaves alternate; stipules subulate, 2-4 × 0.1-0.2 mm; petiole 1-4 mm; leaf blade herbaceous, obliquely elliptic or oblong-oblanceolate, (3-)6-14(-18) × (1.2-)2-5(-6) cm, major lateral veins asymmetric to almost absent, broader outer half auriculate, margin crenate or denticulate, apex acuminate or caudate-acuminate; cystoliths sparse, conspicuous or obscure; nanophyll absent. Staminate inflorescences 0.5-5.5 cm in diam., peduncle 1-8 cm; staminate flowers: tepals 5, elliptic; stamens 5; rudimentary pistil small. Pistillate inflorescences 2-10 mm in diam., peduncle (absent to)1.5-7(-22) mm; pistillate flowers: tepals 5. Achenes ellipsoidal, tuberculate. Fl. Oct-May.
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