Herbs perennial, dioecious. Stems erect or ascending, usually simple, glabrous or pubescent. Leaves alternate; stipules subulate, 2-8 × 0.1-0.3 mm; petiole 3-8 mm; leaf blade herbaceous or papery, obliquely oblong, obliquely lanceolate, or oblanceolate, (4-)8-15 × (1.5-)2.5-5.7 cm, major lateral veins asymmetric to almost absent, broader outer half auriculate, margin crenate or serrate, apex cuspidate, acuminate; cystoliths obscure or conspicuous; nanophyll absent. Staminate inflorescences 1-4.5 cm in diam., peduncle 1-9 cm; staminate flowers: tepals 5, elliptic; stamens 5; rudimentary pistil obscure. Pistillate inflorescences 5-9 mm in diam., peduncle 1-15 mm; pistillate flowers: tepals 4-5. Achenes narrowly ellipsoidal, tuberculate. Fl. Oct-Feb.