Herbs perennial or subshrubs, monoecious or dioecious. Stems ascending or erect, branched or simple, strigose. Leaves alternate; stipules triangular, ca. 10 × 3.5 mm; petiole 0.5-2.0 cm; leaf blade papery, obliquely elliptic, elliptic-ovate, or obovate, 5-13(-15) × 3-6.8(-7.4) cm, major lateral veins symmetric, both basal, broader outer half auriculate, margin entire, apex acuminate or cuspidate; cystoliths slightly conspicuous, dense; nanophylls narrowly ovate, 3-4 mm. Staminate inflorescences 1.5-3.5 cm in diam., peduncle 5.5-12 cm; staminate flowers: tepals 5, elliptic; stamens 5; rudimentary pistil subulate. Pistillate inflorescences 6-8 mm in diam., peduncle 2-5 mm; pistillate flowers: tepals 5. Achenes ovoid or narrowly ovoid, tuberculate. Fl. May-Aug.