Herbs perennial or subshrubs, usually dioecious. Stems ascending or erect, branched or simple, glabrous or pubescent. Leaves alternate; stipules triangular, 12-18 × 3-6 mm; petiole 4-19 cm; leaf blade papery, obliquely elliptic or oblong-obovate, 12.5-20 × 5.8-11 cm, major lateral veins symmetric, both basal, broader outer half auriculate, margin entire, apex acuminate; cystoliths conspicuous, dense; nanophylls ovate or narrowly ovate, 3-4 mm. Staminate inflorescences 2-4 cm in diam., peduncle 3.2-10 cm; staminate flowers: tepals 5, nearly elliptic; stamens 5; rudimentary pistil conic. Pistillate inflorescences 2-3 cm in diam., peduncle 2-10 cm; pistillate flowers: tepals 5. Achenes ovoid, tuberculate. Fl. Dec-Jul.