Herbs perennial, monoecious. Stems erect, branched, glabrous. Leaves alternate; stipules linear or subulate, 1.8-2.8 × 0.2-0.3 mm; petiole 0-1 mm; leaf blade papery, obliquely oblong or obliquely elliptic, 2.4-8 × 1-2 cm, unequal, reduced shoot leaves small, venation pinnate, major lateral veins ill-defined, base obliquely cuneate, margin denticulate, apex caudate-acuminate, or rhombic to rhombic-obovate, 0.6-1.5 × 0.3-0.7 cm, apex apiculate or obtuse; cystoliths conspicuous, dense; nanophyll absent. Staminate inflorescences 2-5 mm in diam., peduncle ca. 1 cm. Pistillate inflorescences 3-5 mm in diam., peduncle 1-4 mm; pistillate flowers: tepals 5. Achenes ellipsoidal, smooth. Fl. Apr-May.
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