Herbs perennial, dioecious. Stems erect, usually simple, glabrous or pilose. Leaves alternate; stipules linear or subulate, 6-9 × 0.5-0.6 mm; petiole 1-7 mm; leaf blade herbaceous, oblanceolate-oblong or obliquely elliptic, 14-25 × 5-8.5 cm, penninerved, broader outer half auriculate, margin crenate, apex acuminate; cystoliths conspicuous; nanophyll absent. Staminate inflorescences 1.5-3 cm in diam., peduncle 2.8-8 cm; staminate flowers: tepals 5, elliptic; stamens 5; rudimentary pistil small. Pistillate inflorescences 0.8-2 cm in diam., peduncle 3-7 mm; pistillate flowers: tepals 5. Achenes ellipsoidal, smooth or striolate. Fl. Sep-Oct.