Herbs perennial, monoecious or dioecious. Stems erect or ascending, usually simple, glabrous or pubescent. Leaves alternate; stipules subulate, 2-7 × 0.1-0.3 mm; petiole 1.5-7 mm; leaf blade papery, obliquely elliptic or oblanceolate, 2-15.5 × 1.5-6.5 cm, major lateral veins asymmetric, one basal, the other arising above base, base obliquely cuneate, margin nearly undulate or undulate-crenate, apex cuspidate or acuminate; cystoliths conspicuous, dense or sparse; nanophyll absent. Staminate inflorescences 0.5-4.5 cm in diam., peduncle 2-7 cm; staminate flowers: tepals 4 or 5, elliptic; stamens 4-5; rudimentary pistil obscure. Pistillate inflorescences 0.4-2 cm in diam., peduncle 0.2-5.5 cm; pistillate flowers: tepals 5. Achenes ellipsoidal, tuberculate. Fl. Apr-Dec.
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