Herbs perennial, usually dioecious. Stems ascending or repent, usually branched, glabrous or puberulent. Leaves alternate; stipules subulate, 1-4 × 0.1-0.2 mm; petiole 1-4 mm; leaf blade herbaceous, obliquely rhombic-ovate or lanceolate, (1.2-)2.4-5(-9) × (0.8-)1-2.5(-4.5) cm, major lateral veins asymmetric, one basal, the other arising above base, broader outer half auriculate, margin dentate, apex acuminate; cystoliths obscure or conspicuous; nanophyll absent. Staminate inflorescences 1-1.5 cm in diam., peduncle 0.5-3.5(-7) cm; staminate flowers: tepals 5, elliptic; stamens 5; rudimentary pistil conic. Pistillate inflorescences 3-5 mm in diam., peduncle 0.3-3(-25) mm; pistillate flowers: tepals 5. Achenes nearly ellipsoidal, tuberculate. Fl. May-Dec.