Herbs perennial, dioecious. Stems ascending, simple, densely crispate-hirtellous. Leaves alternate; stipules subulate, 5-10 × 0.2-0.3 mm; petiole 1-3 mm; leaf blade papery, obliquely ovate, elliptic, or obovate, 1.5-4.5 × 1-3.4 cm, unequal, axils with reduced shoot leaves small, 2.5-7 × 2.5-5 mm, major lateral veins asymmetric, one basal, the other arising above base, broader outer half auriculate, margin crenate, apex acute, obtuse or acutate; cystoliths obscure; nanophyll absent, 2.5-7 × 2.5-5 mm. Staminate inflorescences sessile, with 3-7 flowers; staminate flowers: tepals 4 or 5, oblong-obovate; stamens 4-5; rudimentary pistil small. Pistillate inflorescences 4-6 mm in diam., sessile; pistillate flowers: tepals 4-5. Achenes ellipsoidal, tuberculate. Fl. Oct-Nov.