Herbs perennial or subshrubs, monoecious or dioecious. Stems erect, branched, glabrous. Leaves alternate; stipules subulate, 2-3.5 × 0.1-0.2 mm; petiole 0.4-1.6 cm; leaf blade herbaceous, narrowly oblong or lanceolate, 5-15 × 1.6-5 cm, major lateral veins symmetric, both arising above base, base peltate, obtuse or rounded, margin undulate-crenate or nearly entire, apex acuminate; cystoliths conspicuous, dense; nanophyll absent. Staminate inflorescences 0.5-1 cm in diam., peduncle 0.5-1.8 cm; staminate flowers: tepals 5, elliptic; stamens 5; rudimentary pistil very small. Pistillate inflorescences 3-5 mm in diam., peduncle 1.5-5 mm; pistillate flowers: tepals 5. Achenes narrowly ovoid, tuberculate. Fl. Jun-Sep.