Shrubs or small trees, dioecious, deciduous. Branchlets smooth, usually slightly zigzagged. Stipules small, caducous. Leaf blade margin sharply serrulate, crenate, or subentire. Inflorescence a raceme or panicle of thyrses. Flowers usually 4-or 5-merous. Male flowers: stamens inserted on disk margin; filament subulate; anther subglobose or ellipsoid. Female flowers: disk cup-or ring-shaped; ovary semi-immersed in disk, mostly 2-loculed; ovules 2 per locule, basally attached, erect. Fruit a ± dry berry, subglobose or depressed globose, 2-4-seeded. Seeds subglobose, erect; aril thin.
Shrubs or small trees. Leaves alternate, serrate or entire; stipules present. Flowers bisexual, rarely unisexual, in cymes in axillary thyrses. Sepals 4 or 5, rarely 6-8, valvate or slightly imbricate. Petals 5, rarely 4, valvate or slightly imbricate. Stamens 5, rarely 4 or up to 8, inserted on margin of disc; anthers ±introrse. Ovary partly immersed in disc, mostly 2-locular, with 2 ovules per locule, basally attached, erect. Fruit a berry, 1-4 seeded. Seeds with a thin aril.