Shrubs, monoecious or dioecious, glabrous or hairy, hairs simple; branching not phyllanthoid. Leaves persistent, alternate, simple, all well developed; stipules persistent; blade margins entire. Inflorescences unisexual, fascicles or flowers solitary. Pedicels present. Staminate flowers: sepals 5, distinct; petals 5; nectary extrastaminal, annular, crenate; stamens 5; filaments distinct; connectives not extending beyond anthers; pistillode of 3 distinct segments. Pistillate flowers: sepals persistent, 5, distinct; petals 5; nectary annular, crenate; pistil 3(–4)-carpellate; styles 3(–4), distinct, usually 2-fid to 1/2 length, very rarely 2-fid to base. Fruits capsules. Seeds 2 per locule, rounded-trigonous; seed coat dry, smooth; caruncle usually absent, sometimes present. x = 13.