Phyllanthus amarus Schumach. & Thonn.

Carry me seed (en)

Species

Angiosperms > Malpighiales > Phyllanthaceae > Phyllanthus

Characteristics

Monoecious or rarely dioecious completely glabrous erect or ascending annual herb up to 60 cm. tall, rarely taller, often becoming woody at the base.. Orthotropic stems terete, greenish or reddish. Plagiotropic shoots up to 18 cm. long, although more commonly ± 12 cm. long, the older ones usually co-axillary with secondary ortho-tropic shoots.. Cataphylls linear-lanceolate, 1 mm. long, pale green. Cataphyllary stipules obliquely lanceolate, 2 mm. long, brownish.. Leaves of the plagiotropic shoots distichous, folding up at sundown; petioles 0.3–0.4 mm. long; blades oblong or elliptic-oblong, less commonly elliptic-obovate or elliptic-oblanceolate, 5–12 mm. long, 2–6 mm. wide, rounded to subtruncate at the apex, rounded or rounded-cuneate at the base, membranaceous, lateral nerves 4–6 pairs, looped, inconspicuous above, slightly prominent beneath, bright green above, paler and somewhat glaucous beneath.. Stipules linear-lanceolate, 1 mm. long, cream with a brownish median line.. Male flowers commonly occurring with ♀ in some of the axils in the upper parts of the flowering shoots, ♀ solitary in the remaining axils.. Male flowers: pedicels 0.75 mm. long; sepals 5, ovate, adaxially concave, 0.5 mm. long, acute, pale greenish cream with white margins; disc-glands 5, free, minute, somewhat irregular in outline, flattish, tuberculate; stamens 3, filaments connate to form a column 0.2 mm. high, anthers subsessile, reniform, ± vertically held, 0.25 mm. across, somewhat obliquely dehiscent, buff-coloured.. Female flowers: pedicels 1 mm. long, becoming 1.5 mm. long in fruit, apically thickened; sepals 5, ovate-oblong, 0.75–1 mm. long, 0.3–0.5 mm. wide, subacute, yellowish green or greenish white with white margins; disc flat, thin, 5-lobed, the lobes triangular, with or without 1–2 lateral lobules near the base; ovary subsessile, subglobose, 0.3 mm. diameter, smooth; styles free, minute, 0.1 mm. long, suberect, later ± spreading, shortly bifid, the stigmas rounded, cream.. Fruit depressed-trilobate-subglobose, 1 mm. long, 2 mm. diameter, smooth, ochreous-olivaceous.. Seeds triquetrous, 1 mm. long, with 5–8 well-marked longitudinal ridges on the dorsal facet, and ± 7 less well-marked concentric ridges on each ventral facet, and with innumerable transverse striae between the ridges, uniformly pale greyish-brown.
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Annual herb, glabrous, erect, 1-5 dm high; monoecious; main stem smooth, terete, usually 1.5-2 mm thick; scale-leaves scarious, brownish, 1-2 mm long. Deciduous branchlets 4-12 cm long, ca 0.5 mm thick, subterete (never sharply winged), smooth or proximally roughened, with ca 15-30 leaves. Leaves thin; petioles 0.3-0.5 mm long; stipules lanceolate, 0.8-1.3 mm long; blades elliptic-oblong or somewhat obovate, mostly 5-11 mm long and 3-6 mm broad, rounded to apiculate at the tip, only slightly (if at all) inaequilateral at base, the margins smooth or obscurely roughened. Cymules mostly bisexual, the proximal 1 or 2 axils with staminate flowers, the other axils each with one staminate and one pistillate flower. Staminate flowers with pedicels 0.6-1.3 mm long; calyx-lobes 5 (rarely 6), ovate or elliptic, 0.3-0.6 mm long; disc-segments 5, entire; stamens 3 (rarely 2), the filaments connate into a column 0.2-0.3 mm high, the anthers dehiscing obliquely or horizontally; pollen grains 3-colporate. Pistillate flowers with pedicels becoming 1-2 mm long in fruit; calyx-lobes usually 5, obovate-oblong, acute or apiculate, becoming 0.9-1 mm long; disc flat, deeply 5-lobed, some-times irregular; ovary smooth, the styles free, ascending, shallowly bifid, less than 0.2 mm long. Capsules smooth, stramineous, 1.9-2.1 mm in diam; seeds sharply trigonous, light brown, 0.9-1 mm long, with 5 or 6 straight parallel ribs on the back.
Herbs, annual, monoecious, 1–5 dm; branching phyllanthoid. Stems: main stems terete, not winged, glabrous; ultimate branchlets subterete, not winged, glabrous or slightly scabridulous proximally. Leaves on main stems spiral, scalelike; stipules not auriculate, pale brown. Leaves on ultimate branchlets distichous, well developed; stipules not auriculate, pale green with pale brown margins; blade elliptic-oblong to obovate, 5–11 × 3–6 mm, base obtuse or rounded, apex obtuse to rounded, often apiculate, both surfaces glabrous. Inflorescences cymules or flowers solitary, proximal staminate with 1–2 flowers, distal bisexual with 1 pistillate flower and 1 staminate flower. Pedicels: staminate 0.6–1.3 mm, pistillate spreading in fruit, (1–)1.2–2 mm. Staminate flowers: sepals 5(–6), pale green, flat, 0.3–0.6 mm; nectary extrastaminal, 5 glands; stamens (2–)3, filaments connate throughout. Pistillate flowers: sepals (5–)6, green with broad pale green to white margins, flat, 0.8–1.1 mm, 1-veined; nectary annular, 5(–7)-lobed. Capsules 1.9–2.1 mm diam., smooth. Seeds uniformly brown, 0.9–1 mm, longitudinally ribbed. 2n = 52 (Jamaica).
Female flowers: pedicels c. 1 mm long, extending to 1.5 mm in fruit, apically thickened; sepals 5, 0.75–1 × 0.3–0.5 mm, not or slightly accrescent in fruit, ovate-oblong, subacute, greenish with white margins; disk c. 0.5 mm across, flat, 5-lobed, the lobes triangular, basally lobulate or not; ovary 0.3 mm in diameter, subsessile, subglobose, smooth; styles 3, 0.1 mm long, free, suberect, later ± spreading, 2-lobed.
Male flowers: pedicels 1 mm long; sepals 5, 0.6 × 0.5 mm, ovate, acute, pale greenish-cream; disk glands 5, free, minute, lobulate; stamens 3, filaments connate into a column c. 0.4 mm high, anthers 0.25 mm across, sessile, vertical, 2 bithecate, reniform, obliquely dehiscent with confluent slits, 1 monothecate.
An annual herb. It grows 10-40 cm high. It almost lies along the ground. It is woody near the base. The leaves are in one plain. The flowers hang from one side of the branch. Flowers are separately male and female. The fruit is round and flattened. It splits into three. The seeds have ribs.
Seeds 0.8–1 × 0.6–0.8 × 0.5–0.7 mm, segmentiform, light brown, with 5–7 longitudinal ridges on the dorsal facet, and 4–6 irregularly concentric ridges on each ventral facet, with innumerable faint transverse striae between them.
Leaf blades 5–10 × 2–5 mm, mostly oblong, rounded-subtruncate at apex and base, membranaceous, dull green above, paler beneath; lateral nerves in 4–6 pairs, looped at the apex, inconspicuous above, not prominent beneath.
Scale leaves 1–1.3 mm long, linear-subulate, blackening at the apex; stipules 1.5 × 1 mm, broadly triangular-lanceolate, asymmetrical at the base, dark brown.
Foliage leaves distichous; petioles 0.3–0.5 mm long; stipules 1 mm long, linear-lanceolate, cream-coloured with a brownish midrib.
Male and female flowers often occurring together in the distal axils, female flowers usually solitary in the proximal axils.
A glabrous erect or ascending annual herb up to 75 cm tall, sometimes woody at the base, monoecious or rarely dioecious.
Lateral shoots up to c. 15 cm long, the older ones usually co-axillary with secondary lead shoots.
Fruit 1–1.2 × 1.5–2 mm, oblate, smooth, ochreous-olivaceous.
A glabrous herb with slender subwoody stems
Lead shoots terete.
A weed.
Life form annual
Growth form herb
Growth support free-standing
Foliage retention deciduous
Sexuality
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.3 - 0.5
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color
Blooming months -
Fruit color
Fruiting months -
Nitrogen fixer -
Photosynthetic pathway -

Environment

A weed in villages, gardens, and cultivated fields. Open localities, waste ground, grassy scrub vegetation and dry deciduous forest, usually on humid, sandy soils, at elevations from sea-level up to 1,000 metres.
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A tropical plant. It grows in sunny places. In Fiji it grows near sea level and up to 50 m altitude. It grows in flooded rice fields. In southern China it grows between 100-600 m above sea level. In Yunnan.
Light -
Soil humidity -
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses environmental use food medicinal
Edible fruits leaves
Therapeutic use Diabetes (unspecified), Malaria (unspecified), Venereal (unspecified), Piscicide (unspecified), Digestive (unspecified), Oliguria (unspecified), Diuretic (unspecified), Anti-infective agents, local (unspecified), Anti-inflammatory agents (unspecified), Antipyretics (unspecified), Antiviral agents (unspecified), Appetite stimulants (unspecified), Asthma (unspecified), Astringents (unspecified), Bronchial diseases (unspecified), Colic (unspecified), Diabetes mellitus (unspecified), Diarrhea (unspecified), Diuretics (unspecified), Dysentery (unspecified), Dyspepsia (unspecified), Edema (unspecified), Fever (unspecified), Graves ophthalmopathy (unspecified), Hemostatics (unspecified), Influenza, human (unspecified), Jaundice (unspecified), Kidney diseases (unspecified), Liver diseases (unspecified), Metrorrhagia (unspecified), Scabies (unspecified), Skin diseases (unspecified), Stomach diseases (unspecified), Ulcer (unspecified), Urologic diseases (unspecified), Wounds and injuries (unspecified)
Human toxicity -
Animal toxicity -

Cultivation

Can be grown by seedlings.
Mode seedlings
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Images

Habit

Phyllanthus amarus habit picture by pratiwir pratiwir (cc-by-sa)
Phyllanthus amarus habit picture by R. Tournebize (cc-by-sa)
Phyllanthus amarus habit picture by R. Tournebize (cc-by-sa)

Leaf

Phyllanthus amarus leaf picture by H M (cc-by-sa)
Phyllanthus amarus leaf picture by Ashraf Elsir (cc-by-sa)
Phyllanthus amarus leaf picture by Anindya Chaudhuri (cc-by-sa)

Fruit

Phyllanthus amarus fruit picture by Etienne Noel (cc-by-sa)

Distribution

Phyllanthus amarus world distribution map, present in Aruba, Angola, Åland Islands, Andorra, Benin, Bangladesh, Bahamas, Belize, Bolivia (Plurinational State of), Brazil, Central African Republic, Congo, Cook Islands, Comoros, Costa Rica, Cuba, Cayman Islands, Dominica, Ecuador, Ethiopia, Fiji, Gabon, Guernsey, Ghana, Gambia, Guinea-Bissau, Guatemala, Guyana, Honduras, Haiti, Indonesia, Jamaica, Kenya, Lao People's Democratic Republic, Liberia, Madagascar, Maldives, Myanmar, Mozambique, Mauritania, Nigeria, Nicaragua, Nauru, Pakistan, Panama, Peru, Philippines, Puerto Rico, Paraguay, Sudan, Senegal, Sierra Leone, Suriname, Seychelles, Chad, Togo, Thailand, Tokelau, Tonga, Tuvalu, Taiwan, Province of China, Tanzania, United Republic of, Uganda, Uruguay, United States of America, Venezuela (Bolivarian Republic of), Wallis and Futuna, and Samoa

Identifiers

LSID urn:lsid:ipni.org:names:326408-2
WFO ID wfo-0000270441
COL ID 4H37G
BDTFX ID -
INPN ID 635708
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Phyllanthus amarus Diasperus nanus Phyllanthus nanus Phyllanthus scabrellus Phyllanthus swartzii Phyllanthus amarus var. baronianus Phyllanthus niruri var. amarus Phyllanthus niruri var. scabrellus Phyllanthus amarus subsp. amarus Phyllanthus niruri var. genuinus