Plants 0.2-10 cm, dark green to yellowish, dull to glossy. Stems with short or long stipelike basal region, simple or irregularly pinnate. Stem leaves obscurely seriate, erect when dry, spreading when moist, broadly ovate-acuminate or ovate-lanceolate, not plicate; base auriculate, decurrent, or not; margins plane; costa single, short in proximal leaves, percurrent or short-excurrent in distal leaves; alar cells little differentiated or in several rows along margin, rectangular or quadrate; medial laminal cells sinuate, prorate, walls porose. Branch leaves differentiated from stem leaves, weakly or strongly seriate in 5 rows, spreading when dry or moist, not plicate; apex flat, acuminate; costa percurrent; medial laminal cells sinuate, weakly or strongly prorate. Seta0.5-1.2 cm. Capsule cylindric; operculum rounded or conic, rostrate. Calyptra covering capsule, smooth, hairy.