Erect stems 2-5(-9) cm, not dendroid; sterile stems to 10 cm. Leaves green or yellow-green, crisped and contorted when dry, flat when moist, ovate, elliptic, oblong-elliptic, or rarely obovate, (3-)4-6(-10) mm; base narrowly long-decurrent; margins toothed to near base, smaller leaves sometimes nearly entire, teeth blunt or less commonly sharp, of (1-)2-3(-4) cells; apex rounded, occasionally truncate or obtuse, rarely retuse, cuspidate or apiculate, cusp occasionally toothed; costa percurrent, excurrent, or rarely subpercurrent; medial laminal cells elongate, rarely ± isodiametric (especially near margins), 45-85 µm, less than 1/2 size near margins, in ± longitudinal, occasionally diagonal rows, collenchymatous, sometimes weakly, walls pitted, pits sometimes indistinct or absent; marginal cells linear, sometimes short-linear or rhomboidal, in (2-)3-4 rows. Sexual condition dioicous. Seta 1-3, yellowish or brownish green, 2-4 cm. Capsule pendent, cylindric, 3-4.5 mm, neck not distinct; operculum conic-apiculate. Spores 18-24 µm.