Erect stems 2-5 cm, not dendroid; sterile stems to 12 cm. Leaves green to yellow-green, crisped and contorted when dry, flat when moist, broadly elliptic, ovate, orbicular, or oblong-elliptic, (1-)2-6(-8) mm; base not decurrent or rarely short-decurrent; margins weakly to moderately toothed distally or to 3/4 leaf length, rarely to near base, often entire on sterile stems, teeth blunt, of 1 (or 2) cells; apex broadly rounded, obtuse, or occasionally truncate or retuse, usually narrowly mucronate or cuspidate, cusp bent to side, rarely toothed; costa percurrent or excurrent; medial laminal cells elongate or occasionally ± isodiametric, (30-)50-65(-85) µm, distinctly smaller near margins, often less than 1/2 size, in longitudinal and ± diagonal rows, not to weakly, rarely strongly, collenchymatous, walls pitted, pits sometimes indistinct or absent; marginal cells short-linear or rhomboidal, sometimes linear in larger leaves, in 2-4 rows. Sexual condition dioicous. Seta 1(-3), yellow to reddish brown, 1.8-4.5 cm. Capsule pendent, cylindric or oblong-cylindric, 3-5 mm, neck not distinct; operculum conic-apiculate. Spores 25-30 µm.
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