Erect stems 1-2 cm, not dendroid; sterile stems to 6 cm. Leaves light to dark green, sometimes black with age, crisped and contorted when dry, flat or sometimes weakly undulate in older leaves when moist, broadly elliptic or ovate, 2.5-4(-7) mm; base not or weakly decurrent; margins weakly toothed distally or to near base, occasionally entire, teeth blunt, of 1 (or 2) cells; apex broadly rounded, truncate, or retuse, rarely emarginate, apiculate or rarely cuspidate, cusp smooth; costa percurrent, short-excurrent, or rarely subpercurrent; medial laminal cells short-elongate or ± isodiametric, 22-35(-45) µm, slightly smaller near margins, in indistinct longitudinal or diagonal rows, strongly collenchymatous, walls not pitted; marginal cells linear or sometimes short-linear, in 3-5 rows. Sexual condition synoicous. Seta 1-4, yellow, reddish with age, 1.5-3(-4) cm. Capsule horizontal to pendent, cylindric or oblong-cylindric, 2.5-4.5 mm, neck not distinct; operculum rostrate. Spores 22-33 µm.