Erect stems 1-10 cm, ± dendroid; sterile stems to 10(-15) cm. Leaves dark green or yellow-green, contorted when dry, transversely undulate when moist, oblong-lingulate, lingulate, narrowly ligulate, or rarely narrowly elliptic, 6-10(-14) mm; base long-decurrent; margins toothed to near base, teeth sharp, of 1 (or 2) cells; apex obtuse or rounded, occasionally acute, usually cuspidate, cusp toothed; costa excurrent; medial laminal cells elongate or short-elongate, (12-)15-30(-40) µm, slightly smaller near margins, occasionally in longitudinal rows, rarely in diagonal rows, collenchymatous, walls pitted or not; marginal cells linear, in (2-)3-4 rows. Specialized asexual reproduction by stolonlike stems. Sexual condition dioicous. [Seta 1-4, yellow-green, dark red to blackish with age, 2-4 cm. Capsule pendent, oblong-cylindric, 2.5-3 mm, neck not distinct; operculum rostrate. Spores 30-35 µm].