Plants very small, light to dark green or brownish. Stems freely and irregularly branched; hyalodermis absent, central strand narrow or absent; paraphyllia absent; rhizoids or rhizoid initials below or at abaxial leaf insertion, or axillary, rarely forming tomentum, slightly branched, smooth or granular-papillose; axillary hair distal cells 1 or 2, hyaline. Stem leaves erect-appressed to loosely spreading, sometimes subsecund, lanceolate, not plicate, very small, 0.1-0.3(-0.5) mm; base not decurrent; margins plane, entire to serrulate, limbidia absent; apex acuminate, ; ecostate or costa double, very short, indistinct; alar cells scarcely differentiated to subquadrate, region small, inconspicuous, along basal margins; medial laminal cells subquadrate, oblong-rhombic, or short-rhomboidal; marginal cells 1-stratose. Sexual condition autoicous or dioicous. Capsule erect or variously inclined, oblong-cylindric to oblong-obovoid, symmetric or asymmetric; peristome perfect or somewhat specialized; exostome margins entire or dentate distally; endostome cilia absent or 1-3, rudimentary to well developed. Spores 8-13 µm.