Plants small to large, in thin, intricate mats, yellowish to brownish or darker, glossy. Stems creeping, subpinnate; hyalodermis absent, central strand present, weak; pseudoparaphyllia filamentous or broader. Stem and branch leaves similar, ascending-imbricate to homomallous, ovate-acuminate, not to somewhat plicate; base not decurrent; margins somewhat to strongly recurved, entire to rarely slightly serrulate distally; apex acute to slenderly acuminate; costa double, short, often indistinct; alar cells differentiated, quadrate; laminal cells smooth. Specialized asexual reproduction by dehiscent brood branchlets at branch apices. Sexual condition dioicous; perichaetia small, inconspicuous. Seta reddish. Capsule erect, cylindric to subcylindric, symmetric or asymmetric, not contracted below mouth; annulus sharply differentiated, 2-or 3-seriate; operculum obliquely rostrate; peristome double; exostome teeth with external surface finely papillose to papillose-striolate, smooth at apices; endostome basal membrane low or absent, segments narrowly linear, nearly as long as teeth, jointed, cilia absent. Calyptra naked. Spores round, finely granular.