Plants small to somewhat large, dark green to blackish with yellowish green shoot apices. Stems (fine, prostrate), irregularly branched or subpinnate; hyalodermis absent, central strand present; paraphyllia present, filiform; rhizoids basal or sometimes adventitious distally on decumbent shoots, forming tomentum, slightly branched, smooth; axillary hair distal cells 1 or 2, hyaline. Stem leaves erect-spreading, straight or incurved, cordate-ovate to ovate-lanceolate, not plicate, 0.5-1.5 mm; base not decurrent; margins plane, serrulate distally or to base, sometimes entire, limbidia conspicuous or sometimes inconspicuous, 2-4-stratose, cells incrassate; apex gradually or abruptly acuminate, sometimes apiculate, tips often obtuse; costa single, strong, percurrent to short-excurrent, usually confluent distally with limbidia; alar cells undifferentiated; basal laminal cells rectangular or oblong-hexagonal; distal cells short-rhomboidal to oblong-hexagonal, 6-12 × 15-40 µm. Branch leaves similar, slightly smaller and narrower. Sexual condition autoicous. Capsule horizontal to cernuous, oblong-cylindric, asymmetric, sometimes arcuate; peristome hypnoid; exostome margins dentate; endostome cilia appendiculate. Spores 12-17(-25) µm.
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