Perennials. Stems pilosulous to villous and/or stipitate-glandular. Leaves: petioles 3–8(–12+) cm, sometimes raggedly winged; blades 4–20(–40+) × 2–12(–25+) cm, larger usually deeply pinnately lobed, lobes 5–7. Cypselae 3–4 mm, 3-angled or-ribbed. 2n = 30.