Trees, terrestrial, often with stilt-roots; internodes solid. Scars of the stipules transverse; blade basally attached, palmately incised or entire, venation pinnate to sub palmate. Inflorescences branched (or unbranched and subumbellate), with the flowers ± disperse or (in staminate inflorescences) in terminal, globose heads, ebracteate or sometimes bracteate. Tepals of the staminate flowers (3 or) 4, free or connate; stamens (3 or) 4, free. Perianth of the pistillate flower tubular; stigma (sub)peltate or knob-like. Fruit large (1-2 cm long), dry, enclosed by the enlarged, fleshy perianth, the outer layer of which turning black(ish) at maturity, fruit and perianth constitute a "pseudodrupe", endosperm absent; cotyledons thick.