Plants medium-sized to large, green, brown-green, yellowish, or yellow-brown. Stems unbranched to irregularly pinnate, ± in one plane; hyalodermis absent, central strand present; paraphyllia absent; rhizoids or rhizoid initials on stem or abaxial costa insertion, rarely forming tomentum, slightly branched, smooth; axillary hairs well developed, many, distal cells 1-2(-3), yellow or brownish when young. Stem leaves imbricate, erectopatent, patent, or sometimes spreading, straight or falcate, ovate-lanceolate to very broadly ovate, not plicate, longer than 1 mm; base not decurrent (decurrent in P. trifarium); margins plane, entire, very finely or distinctly denticulate, limbidia absent; apex broadly rounded, apiculate, or acuminate, ; costa single to mid leaf or beyond, or double and short; alar cells ± differentiated, quadrate to long-rectangular, inflated or not, yellow when mature, , region indistinctly delimited, transversely triangular or sometimes in one transverse basal row, large; medial laminal cells linear; marginal cells 1-stratose. Sexual condition dioicous. Capsule horizontal to inclined, cylindric, curved; peristome perfect or nearly so; exostome margins weakly dentate distally; endostome cilia 1-4, well developed, nodose. Spores 10-18 µm.