Plants large, in thick, stiff mats, green, yellow-green, orange-green, or black-green. Stems with branches robust, julaceous, apices curving up; central strand present; paraphyllia many, filamentous, not branched. Leaves appressed to julaceous when dry, erect-spreading when moist, dull, ovate, asymmetric, rarely somewhat falcate, (0.5-) 1-1.2 mm; margins recurved proximally; apex gradually acute to rarely short-acuminate, hair-point absent; costa percurrent, green to yellow-green, somewhat sinuate; alar cells transversely elongate, quadrate, or short-rectangular, region medium-sized; medial laminal cells elongate-rhomboidal, to 40 µm, 3-4:1, opaque, distinctly prorate, walls incrassate, strongly pitted; juxtacostal cells usually shorter than medial cells, walls pitted or not. Capsule inclined to suberect, asymmetric, 0.5-1.2 mm; endostome basal membrane 1/3-1/2 exostome length, segments shorter than exostome, cilia well developed to occasionally rudimentary. Spores (10-)12-18(-20) µm.
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