Plants small, in thin mats, dark green or olive green. Stems with branches slender, not julaceous, apices rarely curving up; central strand present; paraphyllia many, foliose, not branched. Leaves catenulate, weakly incurved when dry, erect-spreading when moist, dull, oblong-ovate, ± symmetric, not falcate, 0.6-0.9 mm; margins plane; apex gradually acute to broadly acuminate, hair-point absent; costa not reaching acumen to subpercurrent, green to orange-green, not sinuate, ; alar cells quadrate to transversely elongate, region large; medial laminal cells quadrate to isodiametric, to 10 µm, 1:1, pellucid, strongly mammillose-papillose over lumen or rarely off-centered, papillae often with apical cell pointed, covering most of lumen, walls incrassate, not pitted; proximal juxtacostal cells usually shorter than medial, walls pitted or not. Capsule unknown.