Trees or shrubs. Leaves simple, alternate, margin entire or crenulate-serrulate, petiolate, penninerved. Flowers dioecious, apetalous; disk absent. Male flowers in axillary racemes or spikes; calyx 4-6-parted; sepals imbricate; stamens 2-4; pistillodes absent. Female flowers solitary, axillary; calyx 5-parted; sepals narrow, imbricate; ovary ovoid or oblong-ovoid, 2-celled; ovules 2 per cell; styles petaloid-dilated, 2-or 3-branched at upper portion, papillose. Drupes indehiscent, long pedicellate, 1-locular, 1-seeded; mesocarp fleshy; pyrene stone-hard. Seed testa membranous or crustaceous; endosperm fleshy.