Inflorescences paniculate, divaricately branched with flowers clustered in numerous dense obovoid-globose heads at the ends of the inflorescence branches; flowers sessile; bracteoles absent.
Leaves mostly cordate at the base, minutely punctate and ± glaucous beneath, strongly pinnately nerved; lateral nerves prominent, numerous, tertiary venation subparallel.
Fruit oblong, ellipsoid or ± globular with a thick cartilaginous pericarp, dehiscent; pericarp thickly woody-crustaceous in mature fruits.
Male flower perianth 3–4-lobed; stamen filaments fused into a slender column; anthers 2–6, adnate to the column about the apical part.
Seed testa somewhat thin; aril laciniate; endosperm fatty, ruminate; cotyledons free almost to the base, suberect.
Female flower perianth as in male; stigma sessile, indistinct.
Evergreen trees or lianes, dioecious or monoecious.