Quercus ilex L.

Holly oak (en), Chêne vert (fr), yeuse (fr)

Species

Angiosperms > Fagales > Fagaceae > Quercus

Characteristics

Evergreen, broad-crowned tree to 20 (–25+) m high, the lower branches often pendulous. Bark hard, ± smooth at first, becoming fissured and tessellated (with small block-like scales), brown to grey. Leaves variable in shape, size and toothing; lamina ovate to lanceolate or elliptic, c. 3–6 (–10) cm long, 1.5–3 (–6) cm wide, cuneate, rounded or sometimes truncate or shallowly cordate at base, entire or occasionally (whole margin to remotely) sharply toothed to spiny (spinulose-dentate, especially leaves of juvenile and vegetative shoots), sometimes wavy margin, whitish-tomentose (downy) when young, the upper surface glabrescent, becoming shiny dark green (often with scattered minute stellate hairs), the lower surface grey-or whitish tomentose (the dense indumentum often becoming greyish or tawny); petiole mostly 0.5–1.5 cm long (to 2 cm long in litt.), tomentose; the leaf and branchlet trichomes minute stellate hairs observed with 10× lens. Monoecious: male flowers in pendulous catkins 3–7 cm long [1.5–2.5 cm long, fide Webb et al. (1988)]; female flowers minute, 2 or 3 on short, erect peduncles in leaf axils. Acorns ovoid to ellipsoid or obloid-ellipsoid, (1.5–) 2–3+ cm long, 1–3 clustered on a short downy peduncle; cupule c. one-third to half (–two-thirds) as long as acorn, with appressed scales.
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Large evergreen tree (to c. 25 m tall in cultivation); bark rough and fissured. Shoots and buds grey-tomentose, without surrounding stipules. Petioles 5-15 mm long, grey-tomentose. Stipules subulate. Lamina 3-8 × 1.5-3.5 cm, usually ovate-oblong or ovate-lanceolate, less commonly broad-elliptic, coriaceous, entire or sometimes spinulose-dentate (lvs of juvenile and vegetative shoots often spinulose-dentate), deep shining green and glabrous above, usually grey-or whitish tomentulose beneath; base broad-cuneate or rounded. ♂ catkins 1.5-2.5 cm long, clustered; rachis villous; bracts densely silky; fls dense along rachis; perianth 1-1.5 mm long, villous outside; stamens 4-8. Frs in clusters of 1-3 fertile ones on a short tomentose peduncle, reaching maturity in first year. Cup 1.3-2 cm diam.; scales appressed, ovate, tomentulose. Acorns mostly 1.5-2 cm long, ovoid, ⅓-⅔ enclosed by cup.
A large tree. It grows to 20-25 m tall and spreads 18-22 m wide. It has a dense rounded crown. It is evergreen. The leaves are alternate. The younger leaves are broad and spiny and older leaves are narrow and do not have teeth. The leaves are glossy and dark green but grey underneath. The male flowers are long and yellow catkins. The acorns are round and small. They can be produced singly or in clusters. They are 2/3 rds enclosed in a scaly cup. The acorns are edible.
Life form perennial
Growth form tree
Growth support free-standing
Foliage retention evergreen
Sexuality monoecy
Pollination anemogamy
Spread dyszoochory endozoochory
Mature width (meter) 4.0 - 8.0
Mature height (meter) 15.0 - 20.0
Root system -
Rooting depth (meter) 4.55
Root diameter (meter) 0.4
Flower color
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway c3

Environment

It grows in Mediterranean climates. It can tolerate salt laden winds. It can grow in poor soils. In Melbourne Botanical Gardens. Temperate. Hobart Botanical Gardens. Kyneton Botanical Gardens. It suits hardiness zones 7-11. Arboretum Tasmania.
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Often growing as the dominant species in arid places, maquis, woods and hills on limestone; at elevations up to 1,800 metres in Europe, but to 2,900 metres in N. Africa.
See Holm Oak (Quercus ilex) in Victorian Resources Online (accessed 5 April 2022) and references therein.
Light 5-8
Soil humidity 3-7
Soil texture 2-5
Soil acidity 3-8
Soil nutriment -
Hardiness (USDA) 7-9

Usage

Often cultivated as an ornamental, shade or street tree. Suitable for tall hedges, including as a coastal windbreak tolerant of maritime conditions. The hardwood has been used in construction and for woodcrafts. The bark is a source of tannin. The acorns are used as fodder for pigs. The tree is favoured in the establishment of truffle orchards. The galls have been used in traditional medicine. See de Rigo & Caudullo (2016), and references therein.
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The acorns can be roasted and eaten or dried and ground into flour for baking. The nuts yield an edible oil.
Uses animal food charcoal construction dye environmental use fodder food fuelwood material medicinal oil ornamental seasoning tanning timber windbreak wood
Edible leaves nuts seeds
Therapeutic use Astringents (bark), Diarrhea (bark), Antioxidants (leaf), Cancer(Intestine) (unspecified), Diuretic (unspecified), Fever (unspecified), Stimulant (unspecified), Astringent (unspecified), Tumor (unspecified)
Human toxicity weak toxic (leaf), weak toxic (bark), weak toxic (fruit)
Animal toxicity toxic (leaf), toxic (bark), toxic (fruit)

Cultivation

Plants are grown from seed. These should preferably be planted where they are to grow. They can also be grown from root suckers.
Mode cuttings graftings seedlings
Germination duration (days) 120 - 365
Germination temperacture (C°) 10
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -12
Optimum temperature (C°) 14 - 32
Size -
Vigor -
Productivity -

Images

Habit

Quercus ilex habit picture by Alain Bigou (cc-by-sa)
Quercus ilex habit picture by Paco Yepes (cc-by-sa)
Quercus ilex habit picture by bennop (cc-by-sa)

Leaf

Quercus ilex leaf picture by Charles Redouly (cc-by-sa)
Quercus ilex leaf picture by alexis joly (cc-by-sa)
Quercus ilex leaf picture by Valeria R (cc-by-sa)

Flower

Quercus ilex flower picture by Simon Da Cunha Belves (cc-by-sa)
Quercus ilex flower picture by Lazo Alfonso (cc-by-sa)
Quercus ilex flower picture by Forns Pau (cc-by-sa)

Fruit

Quercus ilex fruit picture by Marie Dolma (cc-by-sa)
Quercus ilex fruit picture by Frank Noack (cc-by-sa)
Quercus ilex fruit picture by Bernard Boutin (cc-by-sa)

Distribution

Quercus ilex world distribution map, present in Albania, Australia, Switzerland, Cyprus, Czech Republic, Spain, France, Greece, Italy, New Zealand, Portugal, Turks and Caicos Islands, and United States of America

Conservation status

Quercus ilex threat status: Least Concern

Identifiers

LSID urn:lsid:ipni.org:names:296290-1
WFO ID wfo-0000291202
COL ID 793PC
BDTFX ID 54442
INPN ID 116704
Wikipedia (EN) Link
Wikipedia (FR) Link

Synonyms

Quercus ilex f. urceolata Quercus glauca Quercus gracilis Quercus integrifolia Quercus montserratensis Quercus murbeckii Quercus pseud-ilex Quercus sinuata Quercus alpina Quercus crispa Quercus expansa Quercus fagifolia Quercus fordii Quercus gramuntia Quercus ilex Quercus ilicifolia Quercus oblonga Quercus smilax Quercus variifolia Quercus bellota Quercus mixta Quercus marcetii Quercus ilex var. depressa Quercus ilex var. oblongifolia Quercus ilex var. constricta Quercus ilex var. cordifolia Quercus ilex var. crispa Quercus ballota var. grandifolia Quercus ballota var. obovatifolia Quercus ilex var. crispata Quercus ilex var. cylindrocarpa Quercus ilex var. denudata Quercus ilex var. ambigua Quercus ilex var. angustifolia Quercus ilex var. aperta Quercus ilex var. argentata Quercus ilex var. brachycalyx Quercus ilex var. cerasifolia Quercus ilex var. conocarpa Quercus ilex var. elegans Quercus ilex var. ellipticifolia Quercus ilex var. eucalyptoides Quercus ilex var. expansa Quercus ilex var. fagifolia Quercus ilex var. fagifolia Quercus ilex var. fimbriata Quercus ilex var. fordii Quercus ilex var. geminiflora Quercus ilex var. genuina Quercus ilex var. glaucocaesia Quercus ilex var. gracilis Quercus ilex var. gramunita Quercus ilex var. grandifolia Quercus ilex var. integrifolia Quercus ilex var. intermedia Quercus ilex var. lanceolata Quercus ilex var. latifolia Quercus ilex var. laurifolia Quercus ilex var. longifolia Quercus ilex var. macrocarpa Quercus ilex var. macrophylla Quercus ilex var. marcetii Quercus ilex var. microbalana Quercus ilex var. microcarpa Quercus ilex var. mirabilis Quercus ilex subsp. montserratensis Quercus ilex var. myrtifolia Quercus ilex var. obesa Quercus ilex var. obovatifolia Quercus ilex var. oleifolia Quercus ilex var. oosperma Quercus ilex var. operculata Quercus ilex var. ovatiacutifolia Quercus ilex var. ovifolia Quercus ilex var. parvifolia Quercus ilex var. pisiformis Quercus ilex var. plicata Quercus ilex var. polycarpa Quercus ilex var. reducta Quercus ilex var. revoluta Quercus ilex var. revoluta Quercus ilex var. rhododendrifolia Quercus ilex var. rostrata Quercus ilex var. salicifolia Quercus ilex var. serratifolia Quercus ilex var. sinuosifolia Quercus ilex var. smilax Quercus ilex var. spinifolia Quercus ilex var. spinulosa Quercus ilex var. stenocarpa Quercus ilex var. strangulata Quercus ilex var. subocculata Quercus ilex var. subrevoluta Quercus ilex var. subsphaerocarpa Quercus ilex var. turbinata Quercus ilex var. undulata Quercus ilex var. undulata Quercus ilex var. urceolata Quercus smilax var. fordii Quercus ilex subsp. gracilis Quercus ilex var. genabii Quercus ilex subsp. refugiorum Quercus ilex var. refugiorum Quercus castellana Quercus ilex f. ambigua Quercus ilex f. microphylla Quercus ilex f. obesa Quercus ilex f. stenocarpa Quercus ilex f. vulgaris Quercus laurei Quercus rotundifolia f. expansa Quercus ilex var. sphaerocarpa Quercus sempervirens Quercus cyclophylla