Plants medium-sized. Stems creeping, ± regularly 1-pinnate; paraphyllia ± filamentous, not or weakly branched; axillary hairs 3-celled. Stem leaves broadly ovate to cordate-deltoid; margins plane or recurved, crenulate to crenulate-papillose; costa ending near base of acumen; laminal cells ± quadrate, multipapillose on both surfaces. Branch leaves ovate; margins plane to irregularly recurved, crenulate to crenulate-papillose; apex acute or gradually broadly acuminate; costa 4/5 leaf length, covered with cells apically; laminal cells quadrate to isodiametric, multipapillose on both surfaces. Sexual condition autoicous; perichaetial leaves lanceolate, margins crenulate, not ciliate, costa subpercurrent, laminal cells smooth. Seta smooth. Capsule suberect to horizontal, short-cylindric, arcuate; annulus 2-or 3-seriate; operculum stoutly and obliquely apiculate to rostrate; endostome segments not or narrowly perforate, cilia in groups of 1-3.
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