Plants in loose, wiry turfs. Stems robust, often branching, without tomentum, few coarse, red-brown rhizoids at the stem base or occasionally higher and more abundant in robust plants. Cauline leaves 1.8-2.5 mm, at least some leaves with tattered laminae; most apices present, tending to be epapillose; distal surface cells on costa roughened abaxially by distally-projecting papillae at distal ends of cells; basal cells gradually differentiated from distal laminal cells, pale yellow, smooth, 15-17 µm wide; distal laminal cells 7-11 µm wide.