Vines, herbaceous, perennial, climbing; stems to 1.5 m, slender, quadrangular, glabrous and smooth to sparsely aculeolate. Leaves in whorls of 4; petiole (1.5-)2.5-5.5(-13) cm; blade drying thinly papery, adaxially green, abaxially pale green, ovate-cordiform to suborbicular-cordiform, on lateral branches sometimes ovate, (2-)4-7(-12) × (1-)2-5(-6.5) cm, length/ breadth index 1.5-2, glabrous to scaberulous, base cordulate to cordate, margins retrorsely ciliolate or smooth, apex caudate-acuminate, rarely ± obtuse; principal veins 5, palmate. Inflorescence thyrsoid, leafy, with terminal and axillary, few-to many-flowered cymes; axes glabrous and smooth or sparsely aculeolate; bracts linear or lanceolate-linear, 1-3.5 mm; pedicels 1-3 mm. Ovary ca. 1 mm, glabrous. Corolla whitish or pale yellow, subcampanulate, glabrous; tube 0.8-1 mm; lobes spreading and somewhat bent, ovate to lanceolate-triangular, ca. 1.4 mm, caudate. Mericarp berry black at maturity, 4-5 mm in diam. Fl. Jul, fr. Oct-Nov.