Sagittaria guayanensis subsp. lappula (D.Don) Bogin

Subspecies

Angiosperms > Alismatales > Alismataceae > Sagittaria > Sagittaria guayanensis

Characteristics

Laticiferous water plant. Leaves floating, ovate with rounded or emarginate apex, base deeply cordate, basal lobes obtuse, 1.5-6.5 cm long measured along the midrib, greatest width 2.5-10.5 cm; submerged phyllodes sometimes present, linear to lanceolate, shortly petioled, 5-11 by 1/3-1/2 cm; nerves 15-17, prominent on the under-surface, the inner 6-8 directed apically, the others towards the lobes, on the undersurface connected by parallel cross-veins under an angle of c. 70° spaced 1-2 mm, upper surface with veins densely reticulate; petiole ribbed, c. 15-30 cm with air-channels and in sicco prominent septations spaced 2-4 mm; sheath with a broad scarious margin. Peduncles 1-7, rather flaccid, floating, ribbed, with wide air-channels, short-patent hairy towards the apex incl. the raceme 15-40 cm. Racemes c. 1/3 as long as the peduncle, with 2-6 whorls of 2-3 flowers; 1-4 lower whorls with ☿ flowers, the upper ones male. Bracts 3 in each whorl, broad-elliptic, connate, rounded at the top, green with scarious margin, 0.8-2 cm long. Pedicels up to 1 cm, after anthesis thickened and recurved. Sepals broadly triangular-ovate, blunt, keeled, 8-11 by 7-10 mm. Petals obovate to orbicular, very broadly rounded at the apex, delicate, white with sometimes a small purple spot above the yellowish base, 12-15(-18) mm. Stamens in the ☿ flowers 8-12, in an interrupted whorl grouped opposite to the sepals, in the ♂ flowers 6-10, in a whorl, exceeding the rudiments of the pistils, filaments 2-3 mm, much dilated and flattened towards the base, fine-papillose; anthers oblong, 1½ by ¾ mm. Carpels co, elliptic to suborbicular, with a dorsal and a ventral, undulate, membranous crest; style terminal directed inward, c. 1 mm; stigma punctiform. Achenes ∞, 3-4 mm, elliptic, short-stalked, with broad, blunt-spiny crest, spines connected by a thin membrane, c. 1 mm high; lateral ridges without or with minute warts; beak broadly scarious-margined; exocarp with a very fine reticulate relief. Seed brown, 0.5 mm.
More
Stolons usually lacking. Floating leaves broadly ovate to cordate-ovate, 1.2-9 × 1-10 cm, base deeply cordate, apex obtuse; submerged leaves linear or lanceolate, 3-11 cm × 5-10 mm. Inflorescences racemose, 8-40 cm, in 2-6 whorls of 3 flowers; bracts connate toward base. Bisexual flowers usually on lower 1-4 whorls; pedicels 1-1.5 cm, thickened. Male flowers with pedicel 2-5 cm. Sepals not reflexed after anthesis, ovate, 0.5-1.5 cm × 3-15 mm. Petals fugacious, obovate, subequal to sepals. Stamens 6 to numerous; anthers yellow. Achenes obovoid or elliptic, 2-4 × 1.5-3 mm, with cristate wing and oblique adaxial beak. Fl. May-Sep, fr. Jun-Nov. 2n = 22*.
Achenes compressed, 2¼-4 by 1¾-3 mm, wings typically deeply crested, rarely nearly smooth, their faces usually prominently 1-ribbed, the rib occasionally remotely echinate, rarely unadorned.
Life form perennial
Growth form herb
Growth support -
Foliage retention deciduous
Sexuality hermaphrodite
Pollination -
Spread -
Mature width (meter) -
Mature height (meter) 0.5
Root system -
Rooting depth (meter) -
Root diameter (meter) -
Flower color -
Blooming months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Fruit color -
Fruiting months
JanFebMar
AprMayJun
JulAugSep
OctNovDec
Nitrogen fixer -
Photosynthetic pathway -

Environment

Ditches and wet rice-fields, 10-1000 m, often in great quantity, fl. Jan.-Dec. Flowers are closed in the morning, open at noon or somewhat later but never very widely, and close at dusk.
Light -
Soil humidity 10-12
Soil texture -
Soil acidity -
Soil nutriment -
Hardiness (USDA) 9-12

Usage

Uses fodder
Edible -
Therapeutic use -
Human toxicity -
Animal toxicity -

Cultivation

Mode -
Germination duration (days) -
Germination temperacture (C°) -
Germination luminosity -
Germination treatment -
Minimum temperature (C°) -
Optimum temperature (C°) -
Size -
Vigor -
Productivity -

Distribution

Sagittaria guayanensis subsp. lappula world distribution map, present in Afghanistan, Argentina, Bangladesh, China, Ghana, Gambia, Guinea-Bissau, Indonesia, Iceland, Madagascar, Mali, Malaysia, Nigeria, Nepal, Pakistan, Sudan, Senegal, Chad, Thailand, Taiwan, Province of China, and Viet Nam

Identifiers

LSID urn:lsid:ipni.org:names:60444560-2
WFO ID wfo-0000770507
COL ID 5KYX2
BDTFX ID -
INPN ID -
Wikipedia (EN)
Wikipedia (FR)

Synonyms

Sagittaria lappula Sagittaria obtusissima Sagittaria triflora Lophiocarpus cordifolius Lophiocarpus lappula Lophotocarpus formosanus Sagittaria cordifolia Sagittaria pusilla Alisma cristatum Alisma hamiltonianum Sagittaria blumei Alisma lappula Sagittaria guayanensis subsp. lappula Lophotocarpus guayanensis var. lappula Lophotocarpus guayanensis var. madagascariensis