Plants 0.3-6 m. Stems: branches red-brown or brownish, villous or pilose to glabrescent; branchlets densely villous to glabrescent. Leaves: stipules marcescent, rudimentary or foliaceous on early ones, foliaceous on late ones, usually prominent, linear to lanceolate, 2-17 mm, apex acuminate; petiole 4-27 mm, pilose adaxially; largest medial blade oblanceolate or obovate to narrowly elliptic, 43-82 × 12-39 mm, 2.2-4.8 times as long as wide, base cuneate or convex, apex acuminate or convex, abaxial surface very densely to sparsely villous-tomentose or short-silky, hairs straight, wavy, or curved, adaxial long-silky or pilose to glabrescent; proximal blade margins entire; juvenile blade densely villous or long-silky. Catkins: staminate 19-45 × 9-17 mm, flowering branchlet 2-9 mm; pistillate slender to stout, 24-83 × 8-17 mm, flowering branchlet 3-37 mm; floral bract tawny, light brown, or bicolor, 1.5-2.5 mm, apex convex to rounded, hairs wavy. Staminate flowers: abaxial nectary 0.1-0.6 mm, adaxial nectary narrowly oblong, oblong, or square, 0.6-0.9 mm; filaments distinct, glabrous. Pistillate flowers: adaxial nectary sometimes flask-shaped; stipe 0.5-2.8 mm; ovary obclavate or pyriform, densely tomentose, short-silky, or pubescent, beak gradually tapering to styles; ovules 12-22 per ovary; styles connate to distinct 1/2 their lengths, 0.3-1.4 mm; stigmas slenderly to broadly cylindrical, 0.4-0.59-0.8 mm. Capsules 5-9 mm. 2n = 76, 95, 114.