Plants 0.2-2.5 m. Stems: branches yellow-brown or red-brown, villous to glabrescent; branchlets densely villous to glabrescent. Leaves: stipules not marcescent, rudimentary or foliaceous on early and late ones, usually inconspicuous, oblong to elliptic or ovate, 1-2.1-4 mm, apex acute to caudate; petiole 2-9 mm, pilose adaxially; largest medial blade (sometimes hemiamphistomatous), sometimes broadly obovate, 17-63 × 6-28 mm, 1.4-3.5 times as long as wide, base sometimes rounded, rarely subcordate, apex sometimes rounded, abaxial surface pilose, moderately densely villous or long-silky to glabrescent, hairs straight or wavy, adaxial (sometimes dull), pilose or villous to glabrescent; proximal blade margins entire or serrulate; juvenile blade sparsely to densely villous or tomentose. Catkins: staminate 10-48 × 5-14 mm, flowering branchlet 2-25 mm; pistillate subglobose or globose, 18-56(-60 in fruit) × 7-21 mm, flowering branchlet 2-26 mm; floral bract tawny, brown, or bicolor, 2-3 mm, apex rounded, hairs wavy or straight. Staminate flowers: abaxial nectary 0.3-1 mm, adaxial nectary narrowly oblong, oblong, ovate, or flask-shaped, 0.5-1.3 mm; filaments distinct or slightly connate, glabrous, or hairy on proximal 1/2. Pistillate flowers: adaxial nectary 0.4-1.4 mm; stipe 0.3-1.3 mm; ovary pyriform, densely villous or tomentose, beak gradually tapering to styles; ovules 10-18 per ovary; styles connate to distinct 1/2 their lengths or more, 0.7-1.6 mm; stigmas slenderly cylindrical, 0.36-0.48-0.72 mm. Capsules 6-7.5 mm.
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