Trees, 2-15 m. Stems: branches flexible to highly brittle at base, gray-brown to yellow-brown, glabrous or villous; branchlets yellow-brown or red-brown, glabrous, densely villous, velvety, or pilose, nodes hairy. Leaves: stipules rudimentary grading into foliaceous or absent on early ones, usually foliaceous on late ones, apex acute, acuminate, rounded or convex; petiole (shallowly or deeply grooved adaxially, margins sometimes touching, sometimes with basilaminar glands, thickening), 3.5-18 mm, pubescent to glabrescent adaxially; largest medial blade lorate, narrowly oblong, narrowly elliptic, lanceolate, or obovate, 53-190 × 11-35 mm, 2.8-9 times as long as wide, base convex, subcordate, rounded, or cuneate, margins crenate, entire, or finely serrulate, apex acuminate, acute, or caudate, abaxial surface glabrous or pubescent, hairs spreading, white and/or ferruginous, adaxial highly or slightly glossy, glabrous or pubescent, midrib sometimes villous; proximal blade margins entire; juvenile blade glabrous or moderately densely long-silky to pilose abaxially, hairs white and/or ferruginous. Catkins: staminate 31-83 × 7-13 mm, flowering branchlet 2-26 mm; pistillate 28-79 × 6-11 mm, flowering branchlet 3-14 mm; floral bract 1.6-3.4 mm, apex rounded or acute, irregularly toothed or entire, abaxially sparsely to moderately densely hairy proximally or throughout, hairs wavy; pistillate bract deciduous after flowering. Staminate flowers: abaxial nectary 0.4-0.6 mm, adaxial nectary oblong, square, or ovate, 0.3-0.6 mm, nectaries distinct; stamens 3-7; filaments (sometimes basally connate), hairy on proximal 1/2 or basally; anthers 0.4-0.6 mm. Pistillate flowers: adaxial nectary square, 0.2-0.6 mm; stipe 1.4-2.8 mm; ovary pyriform, obturbinate, or ellipsoidal, beak slightly bulged below styles; ovules 12-24 per ovary; styles 0.1-0.2 mm; stigmas 0.2-0.23-0.28 mm. Capsules 3-5.5 mm.
Riparian forests along streams, seepage areas, springs, subalkaline or brackish lakeshores, canyons, ditches; at elevations up to 2,200 metres.
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