Shrubs, 3-7 m, (sometimes forming clones by stem fragmentation). Stems: branches (sometimes ± brittle at base) yellow-gray, yellow-brown, or gray-brown, (sometimes weakly glaucous, with sparkling wax crystals), glabrous; branchlets red-brown or brownish, glabrous or pilose, (inner membranaceous bud-scale layer free, separating from outer layer). Leaves: stipules rudimentary or foliaceous on early ones, foliaceous on late ones, apex acute or rounded; petiole convex to flat, or shallowly grooved adaxially, 4-19 mm, pilose, velvety, or pubescent to glabrescent adaxially; largest medial blade (sometimes amphistomatous), lorate, narrowly elliptic, elliptic, lanceolate, or narrowly oblanceolate, 42-90 × 8-32 mm, 2.8-3.9-5.6 times as long as wide, base rounded, convex, or subcordate, margins flat, entire, serrulate, crenulate, or sinuate, apex acuminate to acute, abaxial surface glaucous, glabrous, pilose, or sparsely long-silky, hairs straight, adaxial dull or slightly glossy, glabrous, pilose, sparsely long-silky, especially midrib; proximal blade margins entire, serrulate, or crenulate; juvenile blade reddish or yellowish green, glabrous or sparsely to moderately densely long-silky throughout, hairs white. Catkins flowering as leaves emerge; staminate stout, slender, or subglobose, 10-45 × 6-12 mm, flowering branchlet 0.5-2 mm; pistillate loosely to densely flowered, stout or subglobose, 13.5-38 × 7-15 mm, flowering branchlet 0.5-7 mm; floral bract brown, tawny, or bicolor, 0.6-1.2 mm, apex acute or rounded, abaxially glabrous or sparsely hairy, hairs curly. Staminate flowers: adaxial nectary narrowly oblong, oblong, square, or flask-shaped, 0.4-0.9 mm; filaments distinct or connate less than 1/2 their lengths, glabrous; anthers yellow or purple turning yellow, (ellipsoid or globose), 0.4-0.8 mm. Pistillate flowers: adaxial nectary oblong, square, or ovate, 0.3-0.9 mm, shorter than stipe; stipe 0.9-3.8 mm; ovary pyriform or ovoid, glabrous, beak gradually tapering to styles; ovules 12-24 per ovary; styles 0.1-0.6 mm; stigmas flat, abaxially non-papillate with rounded tip, or 2 plump lobes, 0.14-0.2-0.3 mm. Capsules 3-5 mm. 2n = 38.
Home