Stems: branches dark red-brown, slightly or highly glossy, glabrous; branchlets red-brown or yellow-brown, puberulent to glabrescent, hairs wavy, curved, straight, or geniculate. Leaves: stipules absent or rudimentary on early ones; petiole convex to flat, or shallowly grooved adaxially, 4-15 mm, pilose or pubescent adaxially; largest medial blade lorate or narrowly oblong, 40-85 × 8-25 mm, 2.9-4.8(-5.7) times as long as wide, base convex or cuneate, margins flat, entire, serrulate, or crenate, apex acute to sometimes acuminate, abaxial surface (pale), glabrous or sparsely pubescent, hairs white and ferruginous, straight, relatively short and stiff, adaxial highly glossy, glabrous, puberulent, or sparsely tomentose, hairs white and ferruginous; proximal blade margins serrulate; juvenile blade reddish. Catkins: staminate (stout or subglobose), 15.5-42 × 9-16 mm, flowering branchlet 1.5-11 mm; pistillate densely flowered, slender, stout, or subglobose, 25-50 × 10-20 mm, flowering branchlet 3-12 mm; floral bract 1.6-3.6 mm, apex rounded to truncate, entire, abaxially hairy throughout or proximally, hairs white and ferruginous, wavy. Staminate flowers: abaxial nectary (0-)0.6-0.9 mm, adaxial nectary oblong or narrowly oblong, 0.5-1 mm, nectaries distinct or connate and cup-shaped; filaments distinct, hairy on proximal 1/2; anthers (purple turning yellow), ellipsoid or shortly cylindrical. Pistillate flowers: (abaxial nectary present), adaxial nectary oblong, 0.4-1 mm, (nectaries distinct); ovary pyriform; ovules 12-16 per ovary; styles connate or distinct 1/2 their lengths, 0.8-1.2 mm; stigmas flat, abaxially non-papillate with rounded tip, or slenderly cylindrical, 0.3-0.6 mm. Capsules 7-11 mm. 2n = ca. 190, ca. 228.
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