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Inflorescences terminal or terminal and lateral with unbranched axes, the node with reduced leaves and stipules which are not modified into bracts surrounding the young inflorescence; true peduncle glabrous or pubescent, elongating.
Calyx tubes entirely fused to each other; limb-tube ± developed, densely pubescent inside, lobes persistent obtuse to triangular with or without deciduous or persistent appendages.
Ovaries 2-locular; placentas discoidal, attached to the middle of the septum; ovules numerous in each locule, spreading in all directions.
Leaves opposite, petiolate, the nerve axils with or without domatia; stipules deltoid, obtuse or slightly notched at apex, ± persistent.
Stamens inserted in the corolla tube, not exserted; anthers basifixed, introrse; filaments short, glabrous.
Style exserted; pollen presenter spindle-shaped, the receptive areas in distal and proximal parts.
Corolla tube funnel-shaped; lobes oblong, imbricate in the bud.
Individual fruits indehiscent, joined to form a fleshy syncarp.
Seeds ovoid or ellipsoid, not winged.
Terminal vegetative buds flattened.
Small to medium sized trees.
Flowers 4–5-merous.
