Plants medium-sized, green, yellowish, or with red secondary pigment. Stems with branch and shoot apices not pencil-like; epidermal cells usually widened to form partial hyalodermis; axillary hair distal cells 1-4, hyaline when young. Stem leaves ovate or ovate-triangular, gradually narrowed to apex, falcate or strongly so, sometimes straight, concave; base not decurrent; margins distinctly denticulate proximally, distally, or both; apex acuminate, ; costa ending 3/5 to nearly entire leaf length; alar region from margins to costa or nearly so; .
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