Plants medium-sized, green, yellowish, or brownish. Stems with branch and shoot apices not pencil-like; epidermal cells not differentiated; axillary hair distal cells 1 or 2, hyaline when young. Stem leaves ovate or ovate-triangular, gradually narrowed to apex, slightly falcate or straight, slightly concave; base broadly long-decurrent; margins denticulate proximally, entire or sparsely denticulate distally; apex broadly acuminate; costa ending 2/3-9/10 leaf length; alar region from margins almost to costa.
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