Plants glabrous throughout. Young branches lacking wings; leaf-bearing branches elongated. Petiole 0.7-3 cm; leaf blade elliptic to ovate, 5-11 × 2-4.5(-6.5) cm, papery to rarely subleathery, not glaucous, venation dark after drying, secondary veins 4-7 on each side of midvein, base cuneate to broadly cuneate, margin denticulate to serrulate, apex shortly acuminate to acuminate. Flowers axillary to fugacious bracts at base of young shoots or axillary to leaves, solitary; peduncle 1.3-4.9 cm (staminate), 2.1-6.5 cm (pistillate). Tepals 6-10, white, yellow, orange, or pink, largest 5-9 × 4.5-7.5 mm. Staminate flowers: stamens 12-40, ± distinct; pollen 6-colpate. Pistillate flowers: carpels 20-45; pseudostyle narrow. Fruit peduncle 3.5-7.5 cm; torus 2-10.5 cm; apocarps red, 3-8.5 × 3-7.5 mm. Seed testa ± smooth, rugulose, or rarely ± tuberculate. Fl. May-Jul, fr. Jul-Nov.