Young branches lacking wings; leaf-bearing branches elongated. Petiole 1-4 cm, glabrous; leaf blade elliptic to ovate, 5-11(-13.5) × 3-7 cm, papery, not glaucous, abaxially glabrous except for pubescent primary and secondary veins, secondary veins (3 or)4 or 5(or 6) on each side of midvein, base cuneate, broadly cuneate, or truncate, margin denticulate to serrulate, apex shortly to long acuminate. Flowers axillary to fugacious bracts at base of young shoots or axillary to leaves, solitary; peduncle 2.8-5.2 cm (staminate), 5-6 cm (pistillate), glabrous. Tepals 7-10, yellow, largest 6.5-10 × 6.5-9 mm, glabrous. Staminate flowers: stamens 14-19, ± distinct; pollen 6-colpate. Pistillate flowers: carpels 36-50; pseudostyle narrow. Fruit peduncle 4.5-10 cm, glabrous; torus 2-15.5 cm; apocarps red, 4-6 × 3-5 mm. Seed testa smooth to ± rugulose. Fl. Jun-Jul, fr. Jul-Aug. 2n = 28*.