Trees', dioecious. Indumentum of solitary, simple hairs; young parts with glandular hairs and more or less sticky resinous. Leaves paripinnate, 2-4-jugate; rachis not winged; no pseudo-stipules. Leaflets subopposite (to alternate); margin entire; domatia absent. Inflorescences in the defoliated part of the branchlets above the leaf-scars, sometimes axillary, fascicles of a few simpe (female) or sparsely branching (male) thyrses, the basal part with scattered, many-flowered fascicles, the upper part spicate. Flowers: sometimes male and female flowers present in the same inflorescence, then one of the two kinds predominating, and the other may be sterile. Sepals 4 or 5 (or 6), connate in lower fourth, slightly valvate but soon apert, the lobes all equal, not petaloid. Petals absent. Disc uninterrupted, more or less patelliform, sinuate, glabrous or sparsely hairy. Stamens (5 or) 6-8 (or 9), long exserted in male flowers; filaments filiform; anthers basifixed in a cleft, dehiscence introrse, glabrous. Pistil (2-or) 3-(or 4-)locular; ovary sparsely to densely pilose and glandular; style subglabrous, with recurved stigmatic lobes; ovules 1 per locule. fruit a hard-crustaceous, dry berry, 1-(or 2-)celled, not winged, either smooth or with patent, simple or branched, strong thorns, glabrescent; inside glabrous. Seeds completely enveloped by an arillode, which is thin-papery and often adhering to the endocarp when dried.
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