Scleranthus L.

German knotgrass (en), Scléranthe des Volcans (fr)

Genus

Angiosperms > Caryophyllales > Caryophyllaceae

Characteristics

Herbs, annual, biennial, or perennial. Taproots slender. Stems erect to prostrate, branched, terete. Leaves connate proximally, sessile; blade 1-veined, subulate to linear, not succulent, apex acute or obtuse. Inflorescences terminal or axillary, lax to dense cymes; bracts paired, foliaceous. Flowers sessile to subsessile; perianth and androecium perigynous; hypanthium urceolate, abruptly expanded distally; sepals 5, distinct, greenish, lanceolate to awl-shaped, 1.5-4 mm, herbaceous, margins whitish, scarious, apex acute to blunt or obtuse; petals absent; nectariferous disc at base of stamens; stamens 2-10, arising from hypanthium rim; filaments distinct; staminodes absent or 5-8, arising from hypanthium rim, filiform; styles 2, capitate, 0.8-1 mm, glabrous proximally; stigmas 2, terminal, minutely papillate (50×). Utricles ovoid, enclosed in persistent, indurate, shallowly or strongly furrowed, sepal-crowned hypanthium and falling with it, the whole constituting the indehiscent “fruit”; carpophore present. Seeds 1, yellowish, globose, not compressed, smooth, marginal wing absent, appendage absent. x = 11 [12].
More
Annual, biennial, or perennial herbs. Stems pubescent on one or two opposite sides, procumbent or erect. Leaves exstipulate, very narrow, joined at the base. Inflorescence mostly reduced dichotomous cymes, or flowers solitary in the leaf axils, at the base enveloped by two scarious bracts. Flowers perigynous; hypanthium urceolate, about as long as the sepals. Sepals 4 or 5, with scarious margin. Petals 0. Stamens 1-10, connate at the base, sometimes alternating with staminodes. Ovary 1-ovuled; styles 2 with 2-or 3-branched stigma. Fruit a membranaceous utricle surrounded by the thickened hard hypanthium. Seeds compressed, glabrous with thin testa.
Fls in cymes or solitary, minute and greenish, hermaphrodite. Sepals 4 or 5 from rim of urceolate perig. cup; petals 0; stamens 1-10; ovary 1-celled with 1basal ovule (occ. 2); styles 2, free to base, stigmas 2; fr. an ind. 1(-2)-seeded nutlet enclosed within hardened perig. cup and persistent sepals; seed smooth, lenticular. Annual to perennial, us. densely branched, herbs with small, subulate, opp., connate, exstipulate lvs. Number of spp. variously estimated at from 10 to 150 in Europe, Asia, Africa, S. America, Australia, New Guinea and N.Z. Relationships of N.Z. and Australian forms need clarifying.
Annual, biennial or perennial herbs. Lvs opposite, exstipulate, linear to subulate, connate by narrowly scarious bases. Fls solitary or paired or clustered, on long axillary peduncles or not pedunculate; bracts small, scarious or leaflike; epicalyx 0. Sepals 4-5, inserted on rim of flask-shaped perigynous zone. Petals 0. Stamens 1, 2 or 5-10, inserted on rim of perigynous zone. Styles 2. Fr. a 1-seeded indehiscent nutlet, with perigynous zone and calyx adhering to fr. and falling with it when ripe. Seeds ovoid or ellipsoid, smooth, not winged.
Fls in compact, terminal and axillary cymose clusters, perfect, perigynous, the cupulate hypanthium resembling a cal-tube; sep 5; pet none; stamens 1–10; ovary ovoid; styles 2, distinct; ovule one on a basal placenta; hypanthium becoming indurate in fr, crowned by the persistent sep and enclosing the membranous utricle; seed obovoid, beaked at the micropylar end; low herbs with diffusely forking stems and opposite, exstipulate lvs connate at base. 10, Old World.
Life form
Growth form herb
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Sexuality hermaphrodite
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Root system tap-root
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Hardiness (USDA) 4-9

Usage

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Cultivation

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Images

Scleranthus unspecified picture

Distribution

Scleranthus world distribution map, present in Australia, New Zealand, Papua New Guinea, and United States of America

Identifiers

LSID urn:lsid:ipni.org:names:19957-1
WFO ID wfo-4000034706
COL ID 63TC2
BDTFX ID 87182
INPN ID 197487
Wikipedia (EN) Link
Wikipedia (FR)

Synonyms

Scleranthus

Lower taxons

Scleranthus uncinatus Scleranthus peruvianus Scleranthus annuus Scleranthus perennis