Plants medium-sized to large (sometimes when submerged), rarely small, not turgid, green, yellow-green, brown, or brown-red. Stems pinnate or sometimes irregularly branched, shoot apices sometimes slightly hooked; hyalodermis well developed, complete. Stem leaves ovate or broadly ovate, gradually or often abruptly narrowed to apex, ± suddenly curved distally, concave, 0.5-1.1 mm wide; apex short-or long-acuminate, often apiculate; costa single, ending beyond mid leaf; alar cells 2-10(-15); medial laminal cells (shorter 1/2 of leaf) 14-95(-120) µm, cell ends square, rounded, or short fusiform-narrowed. Sexual condition dioicous. Capsule with exostome external surface more than 70% dotted basally.
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