Plants terrestrial or epilithic, evergreen, creeping or pendulous from cliffs; upper portion erect or suberect, (15-)30-40(-50) cm, with creeping subterranean rhizome and stolons. Rhizophores restricted to lower part of stem, borne on ventral side in axils of branches. Main stems branched from lower part or from middle upward, pinnately branched, stramineous; unbranched main stem 5-10 cm tall, 1.5-2 mm in diam. in lower part, subquadrangular (carinate in leafy branch portion), sulcate, glabrous; primary leafy branches 6 or 7 pairs, twice pinnately branched, secondary branches forked or once pinnately branched, branchlets dense, adjacent primary branches on main stem 2.4-4.8 cm apart, leafy portion of main stem including leaves 7-8 mm wide at middle, ultimate branches 4-6 mm wide including leaves. Axillary leaves on branches asymmetrical, ovate or triangular, 2-3.2 × 0.9-1.6 mm, base exauriculate, margin denticulate or ciliolate (in lower half). Dorsal leaves asymmetrical, those on main stems obviously larger than those on branches; dorsal leaves on branches contiguous, obliquely ovate, 2.4-3.4 × 1.2-1.8 mm, not carinate, base obliquely cordate, not peltate, margin denticulate or ciliolate (at base), apex acuminate, aristate, or cuspidate. Ventral leaves asymmetrical, those on main stem obviously larger than those on branches; ventral leaves on branches contiguous, slightly ascending, oblong-ovate or oblong, 3.4-4.4 × 1.6-2.2 mm, margin entire, apex acute or obtuse; basiscopic base slightly auriculate, margin entire; acroscopic base enlarged, broader, not overlapping stem and branches, margin denticulate or ciliolate (in basal portion). Strobili solitary, terminal, compact, slightly dorsiventrally complanate, 4-16 × 1.4-2.4 mm; sporophylls dimorphic, slightly dimorphic, or ± uniform, resupinate, not white-margined; dorsal sporophylls broadly orbicular-ovate, slightly carinate, margin shortly ciliolate or denticulate, apex acuminate, with sporophyll-pteryx incomplete and denticulate; ventral sporophylls broadly ovate, carinate, margin denticulate or ciliolate; megasporophylls in basal portion on lower side of strobilus; microsporangia orbicular; microspores pale yellow, megaspores white-yellow.